Author: Spinola, 1808

Type species: Apis truncorum Linnaeus, 1758, designated by Latreille, 1810: 439.
Subgenera: Heriades s.str. Spinola, 1808; Michenerella Krombein, 1950; Rhopaloheriades Griswold and Michener, 1998.
Synonyms:  Eriades Dalla Torre and Friese, 1895; Trypetes Schenck, 1861.
Clade: Anthophila
Family: Megachilidae
SubF: Megachilinae
Tribe: Osmiini
Distinctive traits
  • 2 submarginal cells
  • ventral scopa in females
  • arolium present
  • transverse carina on T1
  • tooth-like axillae
Pictures of distinctive traits.
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Morphologically close genera and how to distinguish them
  • Heriades - Hofferia
Heriades species have tooth-like axillae.
Hofferia species have rounded axillae.
  • Heriades - Stenoheriades
Heriades species have tooth-like axillae and a shorter proboscis. The mandible has three or more teeth.
Stenoheriades species have a long proboscis extending beyond its fossa at rest. The mandible is bidentate.
  • Heriades - Chelostoma, Haetosmia, Hoplitis & Osmia.
Heriades species have a transverse carina on T1 and tooth-like axillae.
Chelostoma, Haetosmia, Hoplitis & Osmia don't have a transverse carina on T1 and have rounded axillae (except for axillae of Osmia (Hoplosmia)).
  • Heriades - Protosmia  
Heriades species have only a vestigial propodeal triangle, but do have a transverse carina on the T1.
Protosmia species have a carinated propodeal triangle. Some don't have a transverse carina on the T1.
General comments on Heriades species identification
Females can be diagnosed by the structure on the clypeus apex and labrum, best seen with open mandibles.
Identification criterion in males are located on the last tergites, sternites and genitalia.
Sorry, but the species identification tool is not yet available for Heriades. 
Please check the reference(s) below for traditional keys.
List of the 6 Heriades species found in Europe (IUCN 2014 + Rasmont et al 2017):
Heriades (Heriades) crenulata Nylander, 1856
Heriades (Heriades) labiata (Pérez, 1896)
Heriades (Heriades) rubicola Pérez, 1890
Heriades (Heriades) truncorum (Linnaeus, 1758)
Heriades (Michenerella) punctulifera Schletterer, 1889
Heriades (Rhopaloheriades) clavicornis Morawitz, 1875
References with identification keys for some of the species:
  • Benoist R. 1929. Les Heriades de la faune française (Hym. Apidae). Annales de la Société entomologique de France, 98: 131-141.
  • Amiet, F., Herrmann, M., Müller, A., & Neumeyer, R. (2004). Apidae 4: Anthidium, Chelostoma, Coelioxys, Dioxys, Heriades, Lithurgus, Megachile, Osmia, Stelis. Fauna Helvetica 9. Centre suisse de cartographie de la faune (CSCF), Neuchâtel 273pp.
  • Falk S.J. (2015) Field Guide to the Bees of Great Britain and Ireland. Bloomsbury wildlife guides. 432 p.

Online resources:
Palearctic Osmiine Bees (Palearctic)
BWARS (UK) 
Exotic Bee ID (World)
Discover Life (World)
WestPalBees (West Palearctic)
Page contributors:
  • Adrien Perrard (Dec. 2019)

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References used to write this page:
  • Michener, C.D. 2007. The Bees of the World, 2nd Edition. The John Hopkins University Press, Baltimore.
  • Michez D., Rasmont P., Terzo, M., Vereecken, N. 2019. Abeilles d'Europes. Hymenoptères d'Europes, Volume 1. N.A.P. Editions.
  • Nieto, A., Roberts, S. P., Kemp, J., Rasmont, P., Kuhlmann, M., García Criado, M., ... & Michez, D. 2014. European red list of bees. Luxembourg: Publication Office of the European Union, 98.
  •  Rasmont, P., Devalez, Jelle, Pauly, A., Michez, D. & Radchenko, V.G. 2017. Addition to the checklist of IUCN European wild bees (Hymenoptera: Apoidea). Annales de la Société entomologique de France 53: 17-32.